mRNA-binding protein expressed during iron starvation; binds to sequence element in 3'-untranslated regions of specific mRNAs to mediate degradation; involved in iron homeostasis; protein increases in abundance and relative distribution to nucleus increases upon DNA replication stress; represses translation in response to iron limitation, downregulating multiple transcripts required for mitochondrial translation and heme biosynthesis; inhibits iron-dependent ribosome recycling factor Rli1p