Kinetochore-associated protein; required for chromosome segregation; required for normal segregation of chromosomes in meiosis and mitosis; recruited to unattached kinetochores by Mps1p-phosphorylated Stu1p where it participates in kinetochore clustering; component of the FEAR regulatory network, which promotes Cdc14p release from the nucleolus during anaphase; potential Cdc28p substrate