Protein that binds tRNA and both methionyl- and glutamyl-tRNA synthetases; forms a complex with methionyl-tRNA synthetase Mes1p (MetRS) and glutamyl-tRNA synthetase Gus1p (GluRS); recruits tRNA to the complex, increases aminoacylation efficiency by stimulating catalysis and ensures cytoplasmic localization of the complex; OB-fold containing protein with unexpected aminoacyl-tRNA deacylase activity; binds quadruplex nucleic acids; protein abundance increases in response to DNA replication stress